Linking together biological models from both the molecular and population levels
has the potential to provide insights that cannot be gained from the study of either
level in isolation.  Computational modeling provides the means to integrate and test
theories which span different temporal and spatial scales, and thus provides an
ideal platform for forging this link.  This paper describes a preliminary model that
links together three key levels of biological theory.  A computational model of gene
regulation is integrated with a model of early land plant morphology, and placed in
an evolutionary environment of competing fitness pressures.  This work provides a
methodological scaffold upon which increasingly complex models at each level of
genotype, phenotype and environment can be built.